Ectoplasm<\/strong><\/h4>\nIt has no trichocysts. It consists of an elastic outer pellicle<\/strong> and a thicker inner layer of contractile fibers or myonemes<\/strong>.<\/p>\na. Pellicle<\/strong><\/p>\n\n- It forms an outer envelope of the bell and also lines the buccal cavity.<\/li>\n
- It is especially thick at the base of the bell.<\/li>\n
- It is devoid of cilia but its basal granules are present in circles showing that its cilia are lost. Basal granules can be demonstrated by the Klein silver-line<\/strong> method.<\/li>\n
- In the stalk, the pellicle is covered by an external cuticle.<\/li>\n<\/ul>\n
b. Myonemes<\/strong><\/p>\n\n- Myonemes form a system of variously running fibers that function differently on contraction.<\/li>\n
- The longitudinal myonemes shorten the body, oblique myonemes pull the disc inwards, and circular myonemes contract and close the peristomial border over the cilia and disc.<\/li>\n
- Myonemes running parallel down the sides of the bell are more visible at the base where they converge before entering the stalk.<\/li>\n<\/ul>\n
Endoplasm<\/strong><\/h4>\nThe granular endoplasm contains nuclei, contractile and food vacuoles, etc.<\/p>\n
a. Nuclei<\/strong><\/p>\n\n- Endoplasm contains a large, elongated,rod-like, and horseshoe-shaped macro-nucleus.<\/strong><\/li>\n
- The macronucleus is highly polyploid with a large amount of chromatin material scattered in the nucleoplasm.<\/li>\n
- A tiny rounded micronucleus<\/strong> occurs in close association with the macronucleus. It is rarely seen in living animals. It can be seen only stain with acetocarmine or any other permanent stains.<\/li>\n<\/ul>\n
b. Contractile vacuoles<\/strong><\/p>\n\n- It is located near the buccal cavity, a clear and permanent pulsating space.<\/li>\n
- It is about 7\u00b5 in diameter when fully distended.<\/li>\n
- It shows diastole 9expansion) and systole(contraction) in a rhythmic manner and pours its contents into an efferent canal, which leads into the buccal cavity.<\/li>\n
- In V<\/em>. picta<\/em> and V<\/em>. monilata<\/em> there are 2 contractile vacuoles.<\/li>\n<\/ul>\n
c. Food vacuoles<\/strong><\/p>\n\n- Many large and small food vacuoles are present in the endoplasm containing food particles.<\/li>\n
- They are formed at the inner end of the cytopharynx and constrict off as rounded spheres into the endoplasm.<\/li>\n<\/ul>\n
d. Cytopyge or cytoproct <\/strong><\/p>\n\n- It is a temporary or permanent opening. The undigested food particles are passed into the buccal cavity through it.<\/li>\n<\/ul>\n
<\/span>Stalk<\/strong><\/span><\/h3>\n\n- The bell-shaped body of Vorticella<\/em> remains attached to the substratum by a long, uniformly, thin, unbranched, and highly contractile stalk.<\/strong><\/li>\n
- It has no cilia.<\/li>\n
- The unbranched stalk distinguishes Vorticella<\/em> from the other ciliates.<\/li>\n
- The older view is that the stalk is formed only by ectoplasm. But, electron microscopy has revealed that the stalk consists of 2 parts, the central canal, and outer sheath.<\/li>\n
- The central canal consists of endoplasm, a contractile cord which is a specialized myoneme, called spasmoneme<\/strong>, and some granules, possibly mitochondria.<\/li>\n
- Spasmoneme is a bundle of spiral fibrils continuous with the myonemes of the bell. It is believed that the contractility of the stalk is due to the contractile ability of spasmoneme.<\/li>\n
- When the stalk is contracted the spiral spasmoneme coils up tightly and looks like spring when viewed under the microscope.<\/li>\n<\/ul>\n
<\/span>Locomotion<\/strong> of Vorticella<\/em> (Bell Animalcule)<\/strong><\/span><\/h2>\n\n- Normally, the cilia of Vorticella<\/em> remain very active and the movement of the stalk are frequent and rapid.<\/li>\n
- When the animal is feeding, its stalk remains fully extended and the bell sways to and fro like a flower in the breeze.<\/li>\n
- Individuals do not move in unison; each individual sways to its own rhythm.<\/li>\n
- But, Vorticella<\/em> is extremely sensitive to any mechanical stimulus. When irritated, all activities cease instantly. The stalk is retracted into a close and delicate spiral, cilia are stilled, the disc is pulled in and closed over by peristomial lip and the rounded body is brought close to the bottom, where it rests motionless until the danger has passed.<\/li>\n<\/ul>\n
<\/span>Nutrition<\/strong> in Vorticella<\/em> (Bell Animalcule)<\/strong><\/span><\/h2>\n\n- The mode of nutrition is holozoic as in Paramecium<\/em>.<\/li>\n
- A water current produced by the adoral cilia of the peristome brings bacteria and other small organic particles into the peristomial groove, from where they are further carried into the buccal cavity.<\/li>\n
- From the buccal cavity, food are carried by undulating membranes to the cytopharynx, movement of food is aided by undulations of the inner rows of cilia.<\/li>\n
- The terminal end of the cytopharynx with food particles is pinched off along with some water into the endoplasm forming food vacuoles<\/strong> one after another.<\/li>\n
- The formation of food vacuoles may be compared with the formation of soap bubbles at the end of a pipe.<\/li>\n
- The food vacuole in the endoplasm moves in an irregular cyclosis (unlike Paramecium<\/em>).<\/li>\n
- Digestion is similar to that of Paramecium and the medium, the medium of food vacuoles is the first alkaline, then acidic, and then it becomes alkaline till absorption.<\/li>\n
- The excess digested food forms refractile glycogen granules stored in the endoplasm.<\/li>\n
- The undigested remains are expelled in the vestibule through the cytoproct.<\/li>\n
- Both food and fecal matters pass through the same passage in Vorticella<\/em>.<\/li>\n<\/ul>\n
<\/span>Respiration, Excretion, and Osmoregulation<\/strong> in Vorticella<\/em> (Bell Animalcule)<\/strong><\/span><\/h2>\n\n- Respiration and excretion take place by diffusion through the general body surface as in Paramecium<\/em> and Amoeba<\/em>.<\/li>\n
- Osmoregulation is performed by the contractile vacuole.<\/strong> A single, large, and pulsating contractile vacuole is present in the endoplasm between the disc and the buccal cavity. It opens into the buccal cavity through a permanent opening and pulsates rhythmically showing diastole and systole phases. At the diastole phase, the excess water from the endoplasm is secreted into it and at the systole phase, the water is expelled into the buccal cavity.<\/li>\n<\/ul>\n
<\/span>Contractility and irritability<\/strong> in Vorticella<\/em> (Bell Animalcule)<\/strong><\/span><\/h2>\n\n- Vorticella<\/em> exhibits a high degree of contractility and irritability.<\/li>\n
- It very readily responds to external stimuli. Even a flash of change in its surrounding water or touch of a minute particle will bring about change in the relative position of its parts.<\/li>\n
- When irritated, the first response is always the stalk, which becomes coiled into a close spiral reduces its size, then the disc is withdrawn and the peristome closes over it. As a result, the body form becomes somewhat globular.<\/li>\n<\/ul>\n
<\/span>Reproduction<\/strong>